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:"Rana okinavana"'' redirects here. That taxon was erroneously thought to refer to the Ryūkyū Brown Frog until recently.'' The Kampira Falls frog, Yaeyama harpist frog or harpist brown frog (''Rana okinavana'') is a species in the true frog family (Ranidae). Until recently known as ''Rana psaltes'', it is found on Ishigaki and Iriomote in the Yaeyama Islands of Japan, as well as on Taiwan.〔Matsui (2007)〕 It is a mid-sized stout brown frog, with a distinctive suprabrachial gland and a fold of skin running from the shoulders to the hips. The latter features unite it with other species of ''Rana'', which may belong into ''Babina'' or ''Nidirana'' if these are considered valid.〔 Its natural habitats are subtropical or tropical moist lowland forests, subtropical or tropical seasonally wet or flooded lowland grassland, swamps, freshwater marshes, and canals and ditches. It is threatened by habitat loss. ==Taxonomy== This frog has a confusing taxonomic history which is entwined with that of the Ryūkyū Brown Frog. Oskar Boettger described a frog species from the Ryūkyū Islands as ''Rana okinavana''.〔Boettger (1895a,b)〕 He could not tell for certain where these specimens were collected as they had reached him via an animal trader and not from the actual collector, but suspected it to be Okinawa in the central Ryūkyūs, the most accessible island of the archipelago.〔 By 1907 it was known that what supposedly was the same medium-sized brown frog also occurred on the Yaeyama Islands in the southern Ryūkyū archipelago,〔Stejneger (1907)〕 though subsequently, Boettger's frog could ''only'' be found there. Later still, the presence of the brown frogs of Okinawa was verified and they were studies, with scientists believing that Boettger's taxon applied to them. Eventually the situation had changed to ''R. okinavana'' supposedly referring to the Okinawan frogs only, with the Yaeyama frogs being either ignored or believed to belong to another species altogether. This was the situation by the time World War II interrupted further research, with some even believing it did not exist at all.〔 In the mid-20th century, the Okinawan and Yaeyama frogs were again united under ''R. okinavana'', but once again without checking Boettger's type specimens. By the 1970s however, it was realized that the southern frogs, though superficially similar in being about the same size and color, differed significantly from those on Okinawa. Initially identified as ''R. adenopleura'', the Yaeyama Harpist Frog was described as a new species, ''Rana psaltes'', in 1985.〔Kuramoto (1985)〕〔 In 1999, upon examination of the lectotype of Boettger's ''R. okinavana'',〔Specimen SMF 5830: Matsui (2007).〕 it was realized that this animal was very similar to ''R. psaltes'', differing from the Okinawan brown frogs just as the recently described Yaeyama frogs did. It furthermore turned out that Boettger's specimens were included in the same batch from the same collector as his specimens of ''Chirixalus eiffingeri''. These specimens must have been collected on the Yaeyama Islands however, as it certainly does not occur on Okinawa. And thus, the frogs formerly known as ''R. psaltes'' - which had also been found on Taiwan in the meantime - turned out to be the same species that Boettger had described, and hence according to ICZN rules, it is now known as ''R. okinavana'', while the Ryūkyū Brown Frog is yet to receive a scientific name. Furthermore, there are some slight differences between Boettger's types of ''R. okinavana'' and Kuramoto's types of ''R. psaltes'' which were collected on Iriomote Island. Should Boettger's specimen, e.g. by ancient DNA analysis, turn out to be from Ishigaki and be subspecifically distinct from the Iriomote population, the latter would become subspecies ''psaltes''.〔 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Kampira Falls frog」の詳細全文を読む スポンサード リンク
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